A fully functional ASIP gene is responsible for the "red", or tan, (phaeomelanin) pigment found in a dogâs coat, when this gene is not functional, black (eumelanin) pigment is produced. In dogs that have a black and tan coat, you will find both functional and nonfunctional ASIP genes in different areas within the coat. The pattern in dogs with black and tan coats can vary from a saddle shape to an almost completely black coat with lighter tan points at the feet or near the shoulders. The spots of tan within the coat are symmetrical on both sides of the body.
ASIP
Multifactorial
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https://pubmed.ncbi.nlm.nih.gov/15965787/ https://vgl.ucdavis.edu/test/agouti-dog https://doi.org/10.1111/age.13154
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